Networks
Bemis, Willy
Professor and John M. Kingsbury Director of Shoals Marine Laboratory
Positions
- Professor, Shoals Marine Laboratory (SML), College of Agriculture and Life Sciences (CALS)
- Academic Director, Shoals Marine Laboratory (SML), College of Agriculture and Life Sciences (CALS)
- Professor, Ecology and Evolutionary Biology (EEB), College of Agriculture and Life Sciences (CALS)
In my role as Kingsbury Director of Shoals Marine Laboratory, I work to provide the best possible undergraduate courses and research internships at our facility on Appledore Island, Maine. With a 40-year track record of educational excellence, SML is working to increase both the breadth and depth of it curriculum to provide todayÕs students with topical and relevant experiences in marine science, sustainability, and conservation. During our 13-week operating season each summer, we offer more than 20 courses for college credit ranging from A Marine Approach to Introductory Biology to Field Marine Science, Diversity of Fishes, Ecology of Animal Behavior, Biological Illustration, Lobster Biology, and Forensics for Marine Biologists. More than 200 undergraduates participate annually in our courses and internships. We also offer adult and family education programs. For additional information and current course offerings, please visit: http://www.sml.cornell.edu/.
Research Areas
- adult and extension education
- aquatic science
- biodiversity
- conservation biology
- earth science
- ecology
- ecosystem biology
- environmental sciences
- evolution
- new life sciences
- science education
- zoology
- Affiliations
- Research
- Publications
- Teaching
- Service
- Background
- Other
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Affiliations
member of graduate field
Research
research overview
- I study the anatomy, development and evolutionary relationships of fossil and living fishes such as sharks, lungfishes, coelacanths, bowfins, sturgeons and paddlefishes.
principal investigator on
- IMPROVING OFF-SEASON RESEARCH ACCESS TO SHOALS MARINE LABORATORY, APPLEDORE ISLAND, MAINE awarded by DIRECTORATE FOR BIOLOGICAL SCIENCES NSF 2012 - 2015
- NEW ENGINE REPLACEMENT FOR THE SHOALS MARINE LAB, VESSEL, J.B. HEISER, UTILIZED FOR THE SHOALS MARINE BIOLOGY PROGRAMS ON APLLEDORE ISLAND, MAINE awarded by ENVIRONMENTAL PROTECT AGENCY 2010 - 2012
- UPGRADE OF SANITATION AND COMMUNICATION SYSTEMS AT SHOALS MARINE LABORATORY awarded by DIRECTORATE FOR BIOLOGICAL SCIENCES NSF 2010 - 2013
area(s) of concentration/expertise
other area(s) of concentration/expertise
- Vertebrate embryology
keywords
- embryology
- evolution
- fish
- molecular evolution
- morphology
- paleontology
- systematics
- vertebrate
submitted impact statement
Publications
individual publications
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academic article
- Social dynamics and dispersal of free-living prairie voles (Microtus ochrogaster).. Journal of Mammalogy. 94:40-49. 2013
- Suckling behaviour in three species of voles. Behavior. 148:551-573. 2011
- Suckling behaviour in three species of voles. . Behaviour. 148:551-573. 2011
- Litter size influences maternal but not paternal care in three species of voles, as measured by nest attendance. Journal of Mammalogy. 88:1420-1426. 2007
- Grouped tooth replacement in the oral jaws of the Tripletail, Lobotes surinamensis (Perciformes: Lobotidae), with a discussion of its proposed relationship to Datnioides. Copeia Journal. 2005:665-672. 2005
- Structure, attachment, replacement and growth of teeth in bluefish, Pomatomus saltatrix (Linnaeus, 1766), a teleost with deeply socketed teeth. Zoology. 108:317-327. 2005
- Methods for preparing dry, partially articulated skeletons of osteichthyans, with notes on making Ridewood dissections of the cranial skeleton. Copeia Journal. 2004:603-609. 2004
- Parental behavior at parturition in prairie voles (Microtus ochrogaster). Journal of Mammalogy. 84:513-523. 2003
- †Protopsephurus liui, a well-preserved primitive paddlefish (Acipenseriformes: Polyodontidae) from the early Cretaceous of China. Journal of Vertebrate Paleontology. 22:209-237. 2002
- Localization and partial characterization of melatonin receptors in amphioxus, hagfish and skate. General and Comparative Endocrinology. 110:67-78. 1998
- An overview of Acipenseriformes. Environmental Biology of Fishes. 48:25-71. 1997
- How many species are there within the genus Acipenser? Environmental Biology of Fishes. Environmental Biology of Fishes. 48:157-163. 1997
- Leo Semenovich Berg and the biology of Acipenseriformes: a dedication. Environmental Biology of Fishes. 48:15-22. 1997
- Letter to Science concerning relationships of coelacanths, lungfishes and tetrapods. Science. 278:370. 1997
- Sturgeon biodiversity and conservation: an introduction. Environmental Biology of Fishes. 48:13-14. 1997
- Sturgeon rivers: an introduction to acipenseriform biogeography and life history. Environmental Biology of Fishes. 48:167-183. 1997
- The threatened status of acipenseriform fishes: a summary. Environmental Biology of Fishes. 48:427-435. 1997
- The Sturgeon’s Plight. Nature. 370:602. 1994
- Early development of the actinopterygian head. I. External development and staging of the paddlefish Polyodon spathula. J. Morphol. 213:47-83. 1992
- Metabolism and ram gill ventilation in juvenile paddlefish, Polyodon spathula (Chondrostei: Polyodontidae). Physiological Zoology. 65:515-539. 1992
- Skin and blood vessels of the snout of the Australian lungfish, Neoceratodus forsteri, and their significance for interpreting the cosmine of Devonian lungfishes. Acta Zoologica (Stockholm). 73:115-139. 1992
- Structure and function of the external gill filaments of embryonic skates (Raja erinacea). Respiration Physiology. 89:1-13. 1992
- Ontogeny of heart function in the little skate, Raja erinacea. Journal of Experimental Biology. 156:387-398. 1991
- Evolutionary morphology of tongue projection in Taricha torosa (Urodela: Salamandridae). Zool. J. Linn. Soc. 99:129-157. 1990
- Convergent evolution of jaw opening muscles in lungfishes and tetrapods. Canadian Journal of Zoology. 65:2814-2817. 1987
- Morphology and function of the feeding apparatus of the lungfish, Lepidosiren paradoxa (Dipnoi) . J. Morphol. 187:81-108. 1986
- Paedomorphosis and the evolution of the Dipnoi. Paleobiology. 10:293-307. 1984
- Structure and growth of lepidosirenid lungfish tooth plates (Pisces: Dipnoi). J. Morphol. 179:73-93. 1984
- Morphology and function of the feeding apparatus in Dermophis mexicanus (Amphibia: Gymnophiona). Zool. J. Linn. Soc. 77:75-96. 1983
- The rostral organ of Latimeria chalumnae: morphological evidence of an electroreceptive function. Copeia Journal. 1982:467-471. 1982
- Kinematics of swimming of penguins at the Detroit Zoo. J. Zool. Lond. 188:411-428. 1979
- Melanin deposits associated with the venom glands of snakes. Journal of Morphology. 155:63-72. 1978
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article
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book
- Functional Vertebrate Anatomy 2001
- A Comprehensive Phylogenetic Study of Amiid Fishes (Amiidae) Based On Comparative Skeletal Anatomy. An Empirical Search for Interconnected Patterns of Natural History. Vol. (supplement to Vol. 18). 1998
- Sturgeon biodiversity and conservation. Vol. 48. 1997
- Osteology and Phylogenetic Relationships of Fossil and Recent Paddlefishes (Polyodontidae) with Comment on the Interrelationships of Acipenseriformes. Vol. supplement to Vol. 11. 1991
- The Biology and Evolution of Lungfishes 1987
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booksection
- Parental Care. IN: Rodent Societies, An Ecological and Evolutionary Perspective 2007
- Parental care. In: Rodent Societies 2007
- Morphology of †Coccolepis bucklandi Agassiz 1843 (Actinopterygii, Coccolepidae) from the Solnhofen lithographic limestone deposits (Upper Jurassic, Germany). IN: Mesozoic Fishes 3 – Systematics, Paleoenvironments and Biodiversity 2004
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chapter
- Occipital structure and the posterior limits of the skull in actinopterygians. Taylor and Francis. 41-62. 2000
- Development of the median fins of the North American paddlefish (Polyodon spathula) and a reevaluation of the lateral fin-fold hypothesis. Mesozoic Fishes II. Systematics and the Fossil Record. 41-68. 1999
- Historical biogeography and historical paleoecology of Amiidae and other halecomorph fishes. Mesozoic Fishes II. Systematics and the Fossil Record. 413-424. 1999
- Interrelationships of Acipenseriformes with comments on “Chondrostei”. Interrelationships of Fishes. 85-115. 1996
- Innervation of the basicranial muscle of Latimeria chalumnae. The Biology of Latimeria chalumnae and the Evolution of Coelacanths. 147-158. 1991
- Skeletal variation in shortnose sturgeon (Acipenser brevirostrum) from the Connecticut River: Implications for the study of fossil fishes. Mesozoic Fishes II. Systematics and the Fossil Record. 69-94. 1991
- Studying physiological evolution: Paradigms and pitfalls. Evolutionary Innovations. Field Museum of Natural History, Spring Systematics Symposium Series. 193-228. 1990
- Feeding systems of living Dipnoi: Anatomy and function. The Biology and Evolution of Lungfishes. 249-275. 1987
- Introduction. The Biology and Evolution of Lungfishes. 3-4. 1987
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report
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review
- Review of: Evolutionary Biology of Primitive Fishes. 114-115. 1986
Teaching
teaching overview
- Willy Bemis teaches courses in vertebrate zoology, marine science, ichthyology, and conservation. During Spring 2008, he offered a seminar on Marine Biodiversity and Conservation for freshman on campus (BIOG 125) and directed or participated in four courses at Shoals Marine Laboratory for SML Summer 2008: FMS, Sharks, A Marine Approach to Introductory Biology, and Boats for Biologists.
teaching activities
- BIOEE-8990: M.S. Thesis Research - Spring 2013
- BIOEE-9990: Ph.D. Dissertation Research - Spring 2013
- BIOEE-8990: M.S. Thesis Research - Fall 2012
- BIOEE-9990: Ph.D. Dissertation Research - Fall 2012
- BIOEE-9990: Ph.D. Dissertation Research - Spring 2012
- BIOEE-9990: Ph.D. Dissertation Research - Fall 2011
Service
outreach overview
- Appledore Island, Maine, is the home of Shoals Marine Laboratory. Each summer, we host hundreds of guests who visit the island and our facilities there. As Director, I am involved in many island tours to explain the natural and human history of Appledore and the Isles of Shoals.
service to the profession
- Fulbright Committee for Graduate Fellowships Committee Member 1988 -
- Sigma Xi Member 1980 -
- Society for Integrative and Comparative Biology Member 1977 -
- American Society of Ichthyologists and Herpetologists Member 1976 -
- National Science Foundation Division of Environmental Biology Served as expert to run panels and manage 50 proposals - 2008
- Research Council, University of Massachusetts Amherst Co-Chair 2002 - 2003
- Research Library Council, University of Massachusetts Amherst Chairperson 2002 - 2003
- Research Council, University of Massachusetts Amherst Member 1999 - 2003
- Research Council, University of Massachusetts Amherst Chairperson - 2003
- Research Council, University of Massachusetts Amherst Chairperson 2001 - 2002
- Research Council, University of Massachusetts Amherst Co-Chair 2000 - 2001
- Personnel Committee, Department of Biology Committee Member 1996 - 1998
- Life Sciences Steering Committee, University of Massachusetts Amherst Committee Member 1995 - 1998
- Society of Integrative and Comparative Biology Program Officer 1995 - 1998
- Life Sciences, Department of Biology, University of Massachusetts Amherst Senior Mentor for Junior Fellows 1993 - 1995
- Biological Sciences Steering Committee Committee Member 1991 - 1995
- Personnel Committee, Department of Biology Committee Member 1992 - 1994
- OEB Admissions Committee Chairperson 1991 - 1994
- OEB Seminar Committee Chairperson 1991 - 1994
- Darwin Postdoctoral Fellowship Program for OEB Founder - 1994
- Division of Vertebrate Morphology, American Society of Zoologists Program Officer 1989 - 1991
- Departmental Honors Program, Department of Zoology, University of Massachusetts Amherst Chairperson 1988 - 1990
- Undergraduate Advising Committee, Department of Zoology, University of Massachusetts Amherst Committee Member 1987 - 1990
- Colloquium Committee, Department of Zoology, University of Massachusetts Amherst Chairperson 1985 - 1987
reviewer or editor for
Background
education and training
- Ph.D. in, University of California - Berkeley 1983
- M.S. in, University of Michigan 1978
- B.A. in, Cornell University 1976
awards and honors
- TEACHnology Fellowship (note: I declined this fellowship because of my relocation to Cornell), 2004
- Distinguished Faculty Award, 2004
- Visiting Scholar, 2002
- Distinguished Faculty Lectureship, 2001
- Samuel F. Conti Faculty Fellowship Award, 2001
- Bass Fellowship, 1991
- Bass Fellow, 1991
- Visiting Researcher, 1990
- NIH National Research Service Award Fellow, 1983
- Chancellor’s Patent Fund Award, 1982
- Annie Alexander Memorial Fellowship, 1981
- NSF Predoctoral Fellowship, 1977
Other
college
- CALS
research keyword
- embryology
- evolution
- fish
- molecular evolution
- morphology
- paleontology
- systematics
- vertebrate
name prefix
- Professor